 
	    LIAS light – Item Descriptions
LIASlight ItemID: 3469
Leptogium papillosum (de Lesd.) P. M. Jørg.
| Characters[*]: | Character States[*]: | 
|---|---|
| global occurrence: | Americas – North America (incl Mexico) | 
| substrate: | bark, cork, plant surface – trunks, branches, twigs | 
| life habit: | lichenized (mutualistic with algal photobionts) | 
| thallus: | foliose (foliaceous), leaf-like | 
| [th] upper surface: | grey(ish) | 
| [th upper surface]: | epruinose | 
| [th marginal and upper surface] specific structures: | present | 
| [th margin] cilia, cilioid structures: | absent | 
| [th upper surface] isidia, isidioid structures: | present | 
| [th upper surface] soredia, soralia, soralioid structures: | absent | 
| [th] morphol substructures (eg areoles, lobes, branches) width [mm]: | (low) 2.0 (high) 5.0 | 
| [th] morphol substructures (eg areoles, squamules): | distantly discontiguous | 
| [th] morphol substructures (eg lobes, branches): | subdichotomous | irregular | 
| [th] lower surface: | grey(ish) | white(ish) | 
| [th lower surface] specific structures: | present | 
| [th lower surface] rhizines, rhizoid structures: | absent | 
| ascomata: | absent | present | 
| ascoma: | apothecial, apothecioid – hymenial | 
| [ascm, if apoth] margin excipular photobionts: | present | abundant | 
| [ascm] paraphyses/-oids: | present | 
| asci: | lecanoralean | 
| [asc] tholus: | thickened | 
| [asc] tholus amyloidity (iodine reaction): | present | 
| [asc] tholus amyloidity pattern: | with amyloid tube, ring (= Collema-, Micarea-, Porpidia-, Psora-types etc) | 
| ascospores: | (median) 8.0 | 
| [asp] shape: | elongate-fusiform, ellipsoidal-fusiform | 
| [asp] length [µm]: | (low) 32.0 (high) 38.0 | 
| [asp] width [µm]: | (low) 9.0 (high) 11.0 | 
| [asp] septa: | present | 
| [asp] septa: | transversely septate | muriformly septate – muriform, submuriform | 
| [asp] transversal septa: | (median) 3.0 | 
| [asp] pigmentation: | hyaline, colourless | 
| [asp] perispore, epispore: | not apparent | 
| secondary metabolites: | absent | 
| primary photobiont: | present | 
| secondary photobionts (eg in cephalodia): | absent | 
| primary photobiont: | cyanobacterial |