 
	    LIAS light – Item Descriptions
LIASlight ItemID: 2977
Lecanora substrobilina Printzen
| Characters[*]: | Character States[*]: | 
|---|---|
| global occurrence: | Americas – North America (incl Mexico) | 
| substrate: | wood – dead, living | 
| life habit: | lichenized (mutualistic with algal photobionts) | 
| thallus: | crustose (crustaceous) – episubstratal – unspecified | 
| thallus: | cracked, fissured, fractured, rimose | compartimentized – areolate, squamulate | 
| [th] upper surface: | grey(ish) | green(ish) brown (olive brown) | yellow(ish) brown | 
| [th upper surface]: | epruinose | 
| [th marginal and upper surface] specific structures: | absent | 
| [th] morphol substructures (eg areoles, squamules): | discontiguous, dispersed (throughout the thallus) | 
| [th] morphol substructures (eg areoles, lobes, branches) upper surface: | verruculose, minutely warted | 
| ascomata: | absent | present | 
| ascoma: | apothecial, apothecioid – hymenial | 
| ascoma [mm]: | (low) 0.55 (high) 1.05 | 
| ascoma: | sessile, superficial | 
| [ascm, if apoth] disc, mazaedium: | plane, flat, flattened, expanded | subconvex, slightly convex | convex | 
| [ascm, if apoth] disc, mazaedium: | green(ish) brown (olive brown) | red(dish) brown (if pale: orange brown) | 
| [ascm, if apoth] disc, mazaedium: | pruinose | 
| [ascm, if apoth] disc, mazaedium pruina: | scarce, sparse, fine | dense | 
| [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: | grey(ish) | green(ish) brown (olive brown) | yellow(ish) brown | 
| [ascm, if apoth] margin excipular photobionts: | present | abundant | 
| [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: | hyaline, colourless | 
| [ascm] paraphyses/-oids: | present | 
| [ascm] paraphyses/-oid cells width [µm]: | (low) 1.0 (high) 1.5 (max) 2.0 | 
| [ascm] paraphyses/-oid apical cells width [µm]: | (low) 1.2 (high) 2.5 | 
| [ascm] epihymenium, epithecium: | brown(ish) yellow (ochraceous, ochre) | 
| asci: | lecanoralean | 
| [asc] tholus: | thickened | 
| [asc] tholus amyloidity (iodine reaction): | present | 
| [asc] tholus amyloidity pattern: | amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc) | 
| ascospores: | (median) 8.0 | 
| [asp] shape: | ellipsoidal | 
| [asp] length [µm]: | (min) 7.0 (low) 7.5 (high) 11.0 (max) 12.0 | 
| [asp] width [µm]: | (min) 3.0 (low) 3.5 (high) 4.5 (max) 5.0 | 
| [asp] septa: | absent – spore lumen unilocular, monolocular | 
| [asp] pigmentation: | hyaline, colourless | 
| [asp] perispore, epispore: | not apparent | 
| secondary metabolites: | present | 
| secondary metabolites: | 4,5-dichloronorlichexanthone | 4-chloronorlichexanthone | arthothelin | norlichexanthone | thiophanic acid | usnic acid | zeorin (hopane-6α,22-diol) | 
| primary photobiont: | present | 
| secondary photobionts (eg in cephalodia): | absent | 
| primary photobiont: | chlorophytaceous – trebouxiaceous, chlorococcoid |